The Sefer Taamei Haminhagim quotes the Eishel Avraham in the name of the Yam Shel Shlomo in Yevamos to hold the Tzitzis between these fingers. He doesn't explain the reason, he just says that there is a very deep "Sod" to doing this. They also quote a Chida in Birchei Yosef who says this is based on Kaballah.
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